The film tells the story of Lucian, the first werewolf born capable of taking human form and the first to be called a Lycan. Viktor, a vampire elder, raises the child, envisioning a race of Lycan slaves guarding the coven's fortress during the day and working as laborers for the vampires at night. The forests and the countryside are filled with savage werewolves born from William's rampage, and human nobles have taken to begging Viktor for protection against the beasts: he grants it in exchange for tributes of silver, which enables him to keep his slaves under control. As Lucian grows up, he and Viktor's daughter Sonja (Rhona Mitra) fall in love, and in their adult years they begin a secretive intimate relationship. Sonja is reckless and insubordinate, and one night Lucian escapes the shackles preventing him from turning werewolf and rescues Sonja from his werewolf brethren. Despite acknowledging that Lucian rescued his daughter, Viktor cannot forgive the escape and has Lucian whipped and imprisoned.